Angiosperms or Flowering Plants

Angiosperm are any of about 300,000 species of the flowering plants, the largest and most diverse group within the kingdom Plantae. Angiosperms represent approximately 80 percent of all the known green plants now living. The angiosperms are vascular seed plants in which the ovule (egg) is fertilized and develops into a seed in an enclosed hollow ovary. The ovary itself is usually enclosed in a flower, that part of the angiospermous plant that contains the male or female reproductive organs or both. Fruits are derived from the maturing floral organs of the angiospermous plant and are therefore characteristic of angiosperms. By contrast, in gymnosperms (e.g., conifers and cycads), the other large group of vascular seed plants, the seeds do not develop enclosed within an ovary but are usually borne exposed on the surfaces of reproductive structures, such as cones.

Unlike such nonvascular plants as the bryophytes, in which all cells in the plant body participate in every function necessary to support, nourish, and extend the plant body (e.g., nutrition, photosynthesis, and cell division), angiosperms have evolved specialized cells and tissues that carry out these functions and have further evolved specialized vascular tissues (xylem and phloem) that translocate the water and nutrients to all areas of the plant body. The specialization of the plant body, which has evolved as an adaptation to a principally terrestrial habitat, includes extensive root systems that anchor the plant and absorb water and minerals from the soil; a stem that supports the growing plant body; and leaves, which are the principal sites of photosynthesis for most angiospermous plants. Another significant evolutionary advancement over the nonvascular and the more primitive vascular plants is the presence of localized regions for plant growth, called meristems and cambia, which extend the length and width of the plant body, respectively. Except under certain conditions, these regions are the only areas in which mitotic cell division takes place in the plant body, although cell differentiation continues to occur over the life of the plant.

The angiosperms dominate Earth's surface and vegetation in more environments, particularly terrestrial habitats, than any other group of plants. As a result, angiosperms are the most important ultimate source of food for birds and mammals, including humans. In addition, the flowering plants are the most economically important group of green plants, serving as a source of pharmaceuticals, fiber products, timber, ornamentals, and other commercial products.


Although the taxonomy of the angiosperms is still incompletely known, the latest classification system incorporates a large body of comparative data derived from studies of DNA sequences. It is known as the Angiosperm Phylogeny Group IV (APG IV) botanical classification system. The angiosperms came to be considered a group at the division level (comparable to the phylum level in animal classification systems) called Anthophyta, though the APG system recognizes only informal groups above the level of order.

General Features

The variety of forms found among angiosperms is greater than that of any other plant group. The size range alone is quite remarkable, from the smallest individual flowering plant, probably the watermeal (Wolffia; Araceae) at less than 2 mm (0.08 inch), to one of the tallest angiosperms, Australia's mountain ash tree (Eucalyptus regnans; Myrtaceae) at about 100 meters (330 feet). Between these two extremes lie angiosperms of almost every size and shape. Examples of this variability include the succulent cacti (Cactaceae), the fragile orchids (Orchidaceae), the baobabs (Adansonia species; Malvaceae), vines, rosette plants such as the dandelion (Asteraceae), and carnivorous plants such as sundews (Drosera; Droseraceae) and the Venus flytrap (Dionaea muscipula; Droseraceae). To understand this vast array of forms, it is necessary to consider the basic structural plan of the angiosperms.

The basic angiosperm form is woody or herbaceous. Woody forms (generally trees and shrubs) are rich in secondary tissues, while herbaceous forms (herbs) rarely have any. Annuals are herbs that complete their growing cycle (growth, flowering, and death) within the same season. Examples of annuals can be found among cultivated garden plants, such as beans (Phaseolus and other genera; Fabaceae), corn (maize, Zea mays; Poaceae), and squashes (Cucurbita; Cucurbitaceae), as well as among the wildflowers, such as some buttercups (Ranunculus; Ranunculaceae) and poppies (Papaver and other genera; Papaveraceae). Biennials are also herbs, but, unlike annuals, their growing cycle spans two years: the vegetative (nonreproductive) plant growth takes place from seed during the first year, and flowers and fruit develop during the second. The beet (Beta vulgaris; Amaranthaceae) and carrot (Daucus carota; Apiaceae) are well-known biennials.

A perennial grows for many years and often flowers annually. In temperate areas the aerial parts of a perennial die back to the ground at the end of each growing season and new shoots are produced the following season from such subterranean parts as bulbs, rhizomes, corms, tubers, and stolons.